The Major and Minor Races of Mankind
Repost from the old site that was shut down. This post is very long and complicated – it runs to 83 pages – but I have tried to make it as easy to understand as possible. Please feel free to dip into it at your leisure. Updated January 28, 2013. Regularly updated.
As you can see by the title, this is an awfully ambitious post. Those who believe that race does not exist, or that Caucasoid, Negroid, Mongoloid and Australoid are outdated terms of no use, might as well bail out right now and save yourself the exasperation.
Recent prior attempts include the usual Mongoloid – Caucasoid – Negroid Three Race Theory, which is discussed below. The main problems with this theory are twofold: that it fails to classify a group called Australoids and that it fails to note the huge split between SE Asians and NE Asians.
From Cavalli-Sforza’s recent work comes an eight-race theory: European Caucasoids, South Asian and North African Caucasoids, Northeast Asian Mongoloids, Southeast Asians extending from Thailand to Indonesia and the Philippines, Pacific Islanders, Australian Aborigines, Negroids and American Indians.
This is not bad, but I would argue that there is no reason to put both Arabs/Berbers and South Indians in one race (see Cavalli-Sforza’s own map below). Genetically, they are quite distant.
From my World Book Encyclopedia 1990 comes a nine-race theory: Negroids, Caucasians, Asians, Polynesians, Micronesians, Melanesians, Aborigines, South Indians and Amerindians. To this I recently added three more very distinct groups, Khoisan (Bushmen), Pygmies and Negritos, to come up with 12 races.
But we can go further than this. If Polynesians and Melanesians are widely regarded as separate races, we should be able to distinguish races based on any other major grouping at least as genetically distant as Polynesians and Melanesians. When I finally found two hapmaps showing the distance between Polynesians and Melanesians, I got the idea for a new race theory based on genetic distance alone.
This theory in most cases is based only on genetic distance, and not physical appearance of physical anthropology. In a few cases, races were grouped into a major group based on appearance – for instance, genetically, Chukchis are in the Caucasian square below, yet they look anything but Caucasian.
Though many distinguish Melanesians and Papuans, Capelli’s (see below) genetic analysis puts them in one race. But see Figures 1-4 below which clearly put them in separate groups. Also, Melanesian and Papuan teeth are very different from each other.
Some people are likely to be upset by this theory.
Surely the Japanese will not be happy to learn that they are virtually identical to the despised Koreans. White Nationalists will not be happy to learn that Turks, Jews, Kurds and Iranians are included in the European race and that they cannot include South Indians with Australoids.
NE Asians and ignorant amateur anthropologists will be unhappy to learn that there is no reason to lump SE Asians with Australoids and that the hated Filipinos (which some refer to as the “niggers of Asia”) are very close to the high-IQ, high-achieving Southern Chinese and the Filipinos haven’t a trace of Negrito in them.
It is standard of NE Asian racialists and amateur anthropologists on the Net to say that the Filipinos are heavily-Negrito.
There are traces of Australoid (Papuan) genes in the Malay, some Indonesians, the Southern Thai and the Coastal Vietnamese, but these admixtures are not large, and the Filipinos haven’t any observable Australoid traces.
Filipinos are closer to Southern Chinese than any other race below, although they are also close to the Aeta Negritos. This is because the Aeta and Ati Negritos are not Australoids genetically but instead are related to SE Asians. Anthropomorphically, they are Australoids.
There is also a more substantial Melanesian component in many Indonesians (except those in Western Indonesia), but there is little if any Australoid, or even Melanesian influence in existing SE Asian populations. It is common amongst Internet anthropologists to lump Melanesians in with Australoids. This is the case anthropomorphically, but not genetically.
In fact, as Figures 1-3 below indicate, they are Asians and are most closely related to other Pacific Islanders. In fact, the distance between SE Asians and Australoids is greater than the distance between NE Asians and Caucasians.
Afrocentrists will be unhappy to learn that various dark folks like South Asians, Melanesians, Papuans and Negritos cannot be considered to be “Black” by any sane definition of the word.
This theory creates nine major races and 113 minor races. It is a work in progress.
Most of this document comes from Cavalli-Sforza’s haplogroup gene map of the human race below.
Figure 1: Cavalli-Sforza’s Principal Coordinate (PC) autosomal DNA haplogroup gene mappings of major human ethnic and racial groups. There are differences between a PC mapping and the tree mappings below.Much of the racial grouping below is based on this map – on genetic distance between groups, not on superficial resemblances between groups. The upper left square can be called NE Asian. The lower left square can be called SE Asian. The upper right square can be called Caucasian. The lower right square can be called African.
Figure 2: Another Cavalli-Sforza map showing general genetic distance, with tremendous overlap with the map above. This map clearly separates out Papuans and Melanesians and also Filipinos and Thais. There is some confusion here regarding the placement of Northern Turkics with Amerindians and whether NW Amerindians should be cleaved off into a separate race.
This map is actually interesting because it implies that there are six major races of humans – not three – NE Asians, SE Asians, Oceanians (Australoids), Pacific Islanders, Caucasians and Africans. As you can see, the distance between NE Asians and SE Asians and between SE Asians and Pacific Islanders is greater than that between NE Asians and Caucasians. SE Asia is clearly an area of profound genetic diversity.
Figure 3: Yet another map, in this case a genetic tree. Once again, Papuans must be cleaved from Melanesians and Thai, and Chinese are clearly separated. This is the first tree that shows the Northern Chinese, and it seems clear it wants to put them with the Koreans and Japanese. This map shows five major races – Caucasians, NE Asians, SE Asians, Africans, Papuans and Aborigines.
Figure 4: More from Cavalli-Sforza showing genetic distance. This was apparently used to map one or both of the maps above. Based on this, I split the Thai off from the Filipinos. This map also shows that Aborigines are most closely related first to Mongolians and Siberians and second to Japanese and Koreans.
I usually wanted about 150 points difference to split off into a separate race, but in some cases I split off closer groups if they were distinguished somewhere else, like in any combination of Figs. 1, 2 or 3. You need to click on it to read it properly.
The initial impulse for this post was this paper in the American Journal of Human Genetics, A Predominantly Indigenous Paternal Heritage for the Austronesian-Speaking Peoples of Insular Southeast Asia and Oceania (Capelli et al 2001). If you look at Table 4 in Capelli, you can see that they carefully delineate out Polynesian and Melanesian groups based on Haplogroup mapping.
Since many scholars of race include both Melanesians and Polynesians as separate races, this table serves to delineate what the proper genetic distance between genetic groups needs to be in order for them to be separate races.
Based on Polynesians and Melanesians as separate races in Table 4 in Capelli, I was able to sort out four more groups in that table, if only to get some idea of the distances between racial groups.
First, an Indonesian Race was separated out, including all but the easternmost island groups such as the Alor that go into Melanesian. Javanese and Sarawak were later included based on Figure 5. Later, based again on Figure 5, the Toraja and Mentawi were separated out, each into their own groups. The Toraja are an ancient farming group in South Sulawesi. The Mentawi are the indigenous peoples of the Mentawi Islands west of Sumatra. They still live a hunter-gatherer lifestyle.
A Lesser Sunda Race was also split out (see Figure 5), but the Alor were not covered, as they lumped more with Melanesians. The Lesser Sunda Race included the Lembata, the Lamaholot, the Manggarai and the Kambera. These people have mixed Indonesian and Melanesian ancestry. The Lembata and Lamaholot live on Lomblen Island east of Flores Island. The Kembara live on Sumba Island and the Manggarai live in the West of Flores Island.
Second, a Filipino-Ami Race, composed of Filipinos and the Ami, a Taiwanese aborigine group (the Filipinos are almost genetically identical to the Ami and are quite close to the Southern Chinese – see Figure 1 in Capelli) was split off.
Third, a South Chinese Race consisting of unknown groups that was later expanded below was split off.
Based on the distances between these clearly differentiated races in Capelli, I was able to plot plot racial distances in Figure 1 above to infer major and minor races based on distance.
All of the groups created via Capelli were then further chopped up based on Cavalli-Sforza here (p. 234-235). An Indonesian Race consisting of Sulawesi, Borneo and Lesser Sunda survived the cut, while the Alor of Lesser Sunda went into Melanesians. Malays themselves are distinct enough to create a Malay race.
The proto-Malay or Temuan, who have some of the most ancient genes on Earth of all of the Out of African peoples, are an ancient aboriginal group in Malaysia. They have an extremely diverse genetic signature (See Figure 5), enough to split off a category all of their own.
The Bidayuh or Land Dayaks are the indigenous peoples of Sarawak. Their genetics are wildly divergent (Figure 5), as we might expect from such an ancient people, hence, they form their own stock.
Some comments are in order.
Although separate NE Asian and SE Asian Major Races were created in order to account for both the vast differences between NE and SE Asians (the distance between NE and SE Asians is greater than the distance between Caucasians and NE Asians) it should still be noted that at a deep level, this is clearly one race.
The Gilyak and Ainu are leftovers from the original Proto-Northeast Asians. The Proto-Northeast Asian homeland was around Lake Baikal maybe 35,000 years ago. The Ainu themselves may go back 18,000 years to the Jomons, who arrived from Thailand. These people resembled Australoids.
In Figure 1 above, Northern Turkic forms a clear race with various Amerindians, yet in Figure 4, they seem to be quite distant. The Buryat have also been linked to Amerindians, even though anthropologically, they are linked to Mongolians and genetically they are close to Koreans.
The North Turkics are closest to the Northern Chinese and the Nepalese, both of which were split off into separate groups. The Manchu and Qiang were added to the Northern Han based on genetics for the Manchu and the fact that the Qiang have an origin in the north. The Yunnan Han, a southern group, oddly cluster with Northern Chinese, as do the Hui.
The Oroqen, a Siberian Tungusic tribe in northeast China that is genetically very divergent, was split off into its own group.
The Nepalese, consisting of Nepalis and Newaris, are genetically Asians, though they resemble Caucasians. They pretty much straddle the line between Caucasians and Asians. A lot of groups close to them – Turkics, Mongols, Northern Chinese, and Altaics, straddle the line between Caucasian and Asian.
Nepalis are closely related to South Indians. They are also close to Central Asians. The Central Asian Race includes the Kirghiz, Karalkalpaks, Uzbeks, Turkmen and possibly others. Although they are mixed Caucasian-Mongoloid people, genetic analysis shows that they can be included with Asians. However, other analysis (Table 2) shows that they are best placed in with Caucasians, though only barely.
Others, such as Kazakhs, are closer to Tuvans and also Mongolians (Table 2). The Kazakhs were placed into a Mongolian Race, somewhat arbitrarily.
The Sherpas were then further split off and placed in with the Yakut (p. 231). All of these splits were based on this data (p. 229). The Tuva were given a separate race based on data showing them splitting away from the Yakut-Sherpas (p. 229)
Northeastern Indians were put into the Mon-Khmer Race somewhat arbitrarily, since this is who they cluster with. There was some confusion. In one paper, the Naga, Apatani, Nishi and Nemang cluster with the Mon-Khmer, and the Adi go in with Tibetans.
The situation is somewhat contradicted by this Y-DNA graph (Reddy 2007), which puts the Apatani, Nishi and Adi, along with the Tripuri, Jamatia, Mog and Chakma, in a single Indian Tibeto-Burman Race. Because of this cluster, and because this group tends to separate somewhat from General Tibetan, I created an Indian Tibeto-Burman Race.
Note that the Tibeto-Burman Tujia, Yizu and Shan cluster away from Indian Tibeto-Burman to some extent. The Mizo and Yizu, Indian Tibeto-Burman groups, cluster more with General Tibetan. However, the Mizo are far enough away from the rest of General Tibetan to warrant their own stock (chart). The Garo also cluster with General Tibetan on Y-DNA, but on Mt-DNA, they are very different (chart) (Reddy 2007).
A group of the Mundas was split off as a Meghalaya Race on the basis of their differentiation on MtDNA (chart) (Reddy 2007). Some Indian Tibeto-Burman groups such as the Bai and the Pnar were included. This race includes the War Jantia, Bhoi, Maram, War Khasi, Kynriam, Nishi, Pnar and Bai. All of these groups are found in Meghalaya or over the border into China.
A group consisting of the Santhal, Naga, Munda, Kurmi and Sudra were split off from this group due to their dramatic difference on MtDNA (chart). This group also lives in NE India.
There is a group of Indo-European speakers in NE India that can be differentiated from the rest of the groups on Mt-DNA. This NE India Indo-European Race consists of the Mahishya, Bagdi, Gaud, Tanti and Lodha.
The Mon-Khmer are close enough to Thai and Southern Chinese in Fig. 4 to be included with the Tai, but they were split off due to the obvious distance in Fig. 1. The Mon-Khmer, Southern Chinese and Thai groups are clearly all closely related.
The Zhuang were split off from Mon-Khmer into a Munda Race on the basis of this autosomal DNA table (p. 235) (Cavalli-Sforza 1994). The She were included because they are close to the Zhuang. The Santhal and Ho were included on the basis of this Y-DNA chart (Reddy 2007). This group is best thought of as an outlier Austroasiatic group.
The Austroasiatic Race consists of the Mon, Zhuang, She, Santhal, Ho and Lyngngam. Most of these groups are found in NE India, but the Mon are in Burma. Most speak Austroasiatic languages, but a some speak Tibeto-Burman or even Indo-European languages. The Nongtrai group with this race in Y-DNA (chart) but not on MtDNA (chart), where they may well form their own group.
The Zhuang are a group in Southern China. They left Central China for Southern China 5000 yrs ago. This group was originally thought to be part of the proto-Tai group in Southern China that later moved down into SE Asia and gave rise not only to the Thai, but also helped form many other SE Asian groups.
At the time of the split from proto-Tai to Tai, the Zhuang went to Guangxi Province and the Tai went to Yunnan. In 1200, the Tai moved down into Indochina and mixed with local groups, becoming the Thai, Lao and Shan.
The Senoi are an ancient group in Malaysia dating back about 4,000-8,000 years. From the close genetic relationship, it seems that the Senoi may have split off from the proto-Zhuang or an earlier group soon after the group left Northern China for Southern China. The Santhal, Ho and Shompen may also have been early split-offs.
The Shompen at least are thought to be a very old group. Originally it was thought that they were remnants of the early people (Negritos) who settled the area, but further research indicated that they are an Austroasiatic group, albeit an ancient one.
Although there is much controversy about the origins of the Senoi (Are they Negritos?) a variety of points of inquiry converge on the notion that they are related to SE Asians.
The Senoi are Veddoids, an ancient group with possible links to the Negritos and the original settlers of Asia 70,000 years ago. There is fascinating evidence for this as Senoi skulls cluster with skulls from the Andaman Islands, Coastal New Guinea and Tamils. Andaman Islanders are Negritos, the New Guinea population is Melanesian and the Tamils are thought to be Veddoid.
The Senoi speak an Austroasiatic language and are also thought to be related to the Vietnamese and the Khmer. Senoi teeth resemble SE Asian and Polynesian teeth. It is thought that the Senoi came down from Southern China and bred in heavily with the Negrito Semang in Malaysia. The Senoi have wavy hair like most Veddoids, though some have straight hair and a few have woolly hair like Negritos.
I recently split the Greater Andamanese and the Onge into two separate major races each based on new data showing that they are profoundly different from all other humans. Whether or not they get separate major races of their own each is open to debate and is determined by the depth of their differences.
However, the data does show that they are each completely separate branches on the human tree. As the Andaman Islanders were the first people to split off after we left Africa and they have been evolving for ~70,000 years in isolation, it figures that they would be extremely different.
I also decided to split Australoids into a macro race alongside Caucasians, Africans and Asians due to charts showing that they are extremely different from all other humans. This group would include for now Papuans, Aborigines and Andaman Islanders.
The Tungus, a group of mostly reindeer-herding tribes, including the Even and the Evenki, were given a separate group based on this map (p. 227). The Evenki are also close to various Tibetan groups, because these Tibetan groups came from NE Asia also.
Amazingly, the Yenisien (of which Ket is the last surviving member) Language Family has now (in 2004) been conclusively tied to the Amerindian Na-Dene Language Family, the first conclusive linking of a New and Old World language family. Even though the Ket presently reside quite a bit to the north of the Altai region where most Amerindians came from, the Ket used to live down near the Altai thousands of years ago.
Northern Turkics include such groups as the Altai, Hazara, Shor, Tofalar, Uighurs, Chelkan, Soyot, Kumandin, Tuva and Teleut. They are located around the Altai Mountains where China, Mongolia and Russia all come together. This is where most of the Amerindians came from.
Evidence for including the Hazara, who speak a language related to Persian, in the Northern Turkic group is a chart that shows the Hazara clustering with the Uighur.
Malay Negritos (the Semang) were given a separate race based on a recent study finding them highly differentiated from other Asian populations. The Jehai and Kensui are related Negrito groups in Malaysia (Figure 5).
Though Cavalli-Sforza includes Berbers barely into the African square, I include them with Caucasians due to their greater resemblance to Caucasians than African, and also due to genetic analyzes that show that they have little Black in them. However, some Berbers are clearly African. Analyses of the more-Caucasian Berbers find that, across the board, they are on average 12% Black.
Tuaregs were given separate races because they are clearly separate from Berbers and all of the African groups in Fig. 1.
However, Tuaregs do cluster (p. 169) with Algerians and Bejas. Since Algerians are Caucasian and most Tuaregs are Africans (though they vary considerably), I had to separate them into major races based on appearance. This is one of those cases where genes flies in the face of physical anthropology.
Bejas are a mixed-race people living in northeastern Africa and speaking a Cushitic language. They look like Ethiopians. Ethiopians are about 57% African and 43% Caucasian – Amhara are 57%, Cushitic are 56% and Tigreans are 53% Black. Since the Beja are a Cushitic group, on that basis, I put the Beja into Africans.
Similarly, Nubians are grouped (p. 169) in with the Caucasian Berbers, although most people consider them to be Black people. With examples like this, you can see why Fig. 1 has Berbers on the border of African and Caucasian.
Figure 1 also puts the Chukchi in the Caucasian square, though they clearly resemble Asians. I lump them in with Asians due to their obvious resemblance to Asians. I included Aleuts with Chukchis due to a recent paper showing a linkage.
Siberian Eskimos were included for the same reason. The entire group was called the Beringian Race. The Koryaks were split into a separate group due to Cavalli-Sforza’s data. The Itelmen were later added to the Koryaks due to evidence showing that they are related. Both were combined into a Paleosiberian Race. The Reindeer Chukchi, apparently a more Siberian group, was split off due to its great (p. 228) genetic distance from other groups.
The Uralic Race was split into a Siberian Uralic Race including the Samoyed, Ket and Nentsy subgroups (p. 227). The Nganasan are an outlier (p. 229) in this group, and there was barely enough evidence to split them into a separate group.
Northern Na-Dene speakers were split from the North American Eskimos whom they resemble (p. 323), on the basis of this tree (p. 227). Similarly, Ge and Tucanoan (linguistic groups) Amerindians were split off from the rest due to great distance (p. 322) between them and the others.
A Fuegian Amerindian Race was created based on evidence that they exhibit extreme genetic differences with all other Amerindians. They are probably the ancestors of the original peopling of the Americas.
The Nootka, or Nuuchahnulth, were also split off due to the finding of a fifth major haplogroup lineage (p. 1166) in them in addition to the main four lineages – A-D – usually found in Amerindians. This line links back to ancient Amerindian remains and goes back to Mongolia.
I started out with a General Amerindian Race, but I decided to split it into four races – Northwest American, Northern, Central and Southern, based on Figure 2. It is true that I could not make these splits on the basis of Figure 1 or the genetic distance charts, but as most serious splits on Figure 2 went into separate races, I decided to split the Amerinds in the same manner.
Further, the Amerinds have some of the greatest internal genetic distances of any geographical group, far more, for instance, than the Europeans and Iranians, so the splitting seemed valid.
South Indians are included with Caucasians based on a general consensus that these are an ancient group of Caucasians. The reason being their resemblance in facial and body structure to Caucasians. In addition, Figure 1 clearly puts them in the Caucasian square, and the other three figures clearly show that they are most closely related to Caucasians.
Although genetic studies say that South Indians are all one race and there is good reason to believe this, Figure 1 delineates South Indians and North Indians into separate groups, though there is a clear transition from one to the other. Figures 2 and 3 reiterate the distinction between South and North Indians.
There is data linking Vietnamese genetically with Cantonese. Vietnamese genetics are very complex and it is all being worked out. They are clearly an Austronesian-Tai mix with heavy S. Chinese admixture and some undetermined amount of Khmer and Cham mixed in. Vietnamese does not include the Montagnards, who are the indigenous people and seem to be related to Negritos.
There is good evidence also linking the Vietnamese and related groups to the Tai, however, there seems to be better evidence linking to them to a small group of mostly Mon-Khmer speakers. The Deang or Paluang, the Jinuo and the Blang lump together with the Vietnamese (Lĭ 2006). The Mon-Khmer speaking Deang live in Yunnan, Burma and Thailand, the Tibeto-Burman speaking Jinuo live in Yunnan and the Blang also live in Yunnan. So the closest living relatives to the Vietnamese people are in Yunnan, and next in Burma and Thailand.
Since there is quite a bit more distance between Filipinos and Thais than between Filipinos and Southern Chinese, I split off Thais into a separate race. I also kept the Filipino-Ami Race above, but added the Guangdong Han (Guangdonren in Chinese) to the group based on evidence that they are linked to the Ami.
Based on Fig. 5, I further refined the Filipino portion of this group into Tagalog, Visaya and Ilocano speakers, while splitting off the Manobo into a separate group, as they are divergent (Fig. 5). Tagalogs are an ethnic group who live mostly in Luzon and Oriental Mindoro, while Visayan languages are spoken in the Visayas region in the central Philippines, encompassing the islands of Panay, Negros, Cebu, Bohol, Leyte, Samar and Palawan. Ilocano speakers are located in the far north of Luzon.
A race called the Southeast China Race was created based on a tight clustering of the Minnan Nan, Hakka, and overseas Chinese of Singapore and Thailand. Based on Figure 5, the Cantonese Han (outside of Hong Kong) were added to this race.
A separate Taiwanese Aborigine Race was split off, based on Cavalli-Sforza’s work. This group, best seen as the principal Taiwanese Aborigine Race, consists of the Atayal, Bunun and Yami. Another Taiwanese Aborigine group, the Paiwan, was split into an Island SE Asian Race based on Cavalli-Sforza. Interestingly, the Paiwan, Atayal and Yami are also somewhat close to the Tai Race (see below).
The Taiwanese Aborigines have an interesting background, and their prehistory is in need of further research.
In addition to the Thais proper, I also include other Tai groups such as the Tai Lue, Tai Kern, Tai Yong and Tai Yuan on the basis of Figure 5. All are found in Thailand. Many groups are related to the Thais. They are the Lao, Shan, Dai, Lahu, Aini and Naxi. The Lahu, Dai and Aini were included on the basis of this report. All of them are found in Yunnan. This group is found in Southern China (especially Yunnan), Laos, Vietnam, Thailand and Burma. The Buyei are also related to the Thai.
Two aboriginal groups of Thailand are so different as to warrant a separate stock each.
The Htin, or Mal, are ancient aborigines of Thailand speaking a Khmuic language. In Figure 5, they are different enough to constitute their own stock.
The Mlabri are a very strange group of hunter-gatherers in Thailand who are very poorly understood. They live very primitive lives. Their genetics is wildly diverse and suggests that they were founded from a small stock only 800 years ago or so. That is, they went through a genetic bottleneck. Some think that they are former farmers who went back to land for some reason. They are one of the most genetically wildly diverse people in Asia (see Figure 5).
Although Fig. 4 suggests that Southern Chinese and the Thai should be grouped together, Figs. 1-3 suggest otherwise. Clearly, the two groups are very close, but I decided to break Southern Chinese off due to the other figures above, especially Figure 1, that suggest they are a separate grouping.
I lumped a number of groups into a Southern Chinese Race, including the Dong, Yi and the Han living in Henan Province, China, based on evidence that they form a group with the Southern Chinese. These groups are found in the Southern Chinese provinces, including Henan, Guangxi, Sichuan, Guizhou, Hainan and Fujian.
I created a Hmong-Mien Race for the Hmong and the Mien, since, while they are close to the Southern Chinese Race, they are different enough to merit their own category (see Figure 5).
Figure 5: Click to enlarge. A good chart of many of the Asian races, showing how well genes and language line up.
The Li is a genetically divergent Chinese ethnic group that forms it’s own outlier between the Southern and Northern Chinese. However, it trends more towards Southern Chinese. They also link up very closely to the Khmer. The suggestion here is that the ancestors of the Khmer were the Li.
What we are learning about Negritos is that instead of forming a distant group, they are often closest to the people they are living around. So the Philippine Negritos (Aeta) are closest to other Filipinos, and the Veddas are closest to other South Asians.
The Mamanwa, a Negrito group on Mindanao Island in the Philippines, are highly divergent from the rest of the Philippine Negritos. The Mamanwa are thought to be remnants of the original Negrito population in the Philippines.
The Palau, a Micronesian group, curiously cluster with Aeta and Agta Negritos, indicating that they may be the remains of the original settlers of SE Asia. The Agta and Aeta cluster together also (Fig. 5). The Aeta and Agta Negritos both live in mountainous areas of Luzon.
The Iraya Mangyans of the Philippines are also quite different, but they are close to the Ati Negritos, also of the Philippines (Fig. 5). The Ati live on Panay Island, in the Visayas Group. The Iraya are a Mangyan group living on Mindoro Island. The Mangyans are not Negritos, but they are still an indigenous group in the Philippines and are different from most Filipinos.
The Toba Batak, a tribe in northern Sumatra, curiously clusters with the Kanaka and Yap Micronesians. On Figure 5, the Karo Batak line up with the Toba Batak. They may be leftovers of the original Melanesian-Polynesian mix that populated Micronesia. The Kanaka is an old name for a Micronesian tribe that lives primarily in the Carolines and the Marshall Islands in the Pacific.
The Veddas are clearly related to the Negritos as one of the sole remaining leftovers of the group that left Africa 70,000 years ago and populated all of Asia. There are interesting links between them and the Toala of Southern Sulawesi and the Senoi of Malaysia. Nevertheless, almost all Veddas except the Kerala Kadar cluster with the South Indian Race.
North Indians include the Punjabis, Central Indic, Punjabi Brahmins, Rajputs, Vania Soni, Mumbai Brahmins, Jats, Kerala Brahmins, Pakistanis and Koli.
South Indians include the Munda, Bhil, Maratha, Rajbanshi, Oraon, Parji, Kolami-Naiki, Chenchu-Reddi, Konda, Kolya, West Bengal Brahmins, Parsi and Gonds. Although many of these groups are thought to be related to Veddas or Negritos and part of the original people of India, they now resemble other South Indians.
Kerala Kadar are a highly diverse Vedda group who are probably the ancestors of the original people of India. They live in the forests of Kerala and resemble Australoids.
The Gurkha and Tharu are two highly diverse groups in Nepal. In Figure 5, the Ladakhi are close to them, so a Himalayan Race was created to encompass them.
The Kanet live in Himachal Pradesh and Gujarat and probably have some Tibetan mixture. The inclusion of the Uttar Pradesh Brahmin with these people in unexplained.
The Nicobarese and the Senoi cluster with the Munda Race on Y-DNA, but on Mt-DNA, they are extremely different (chart here) (Reddy 2007), which is suggested by their ancient origins. Each got a separate race due to their extreme divergence.
The Khoisan were divided into three groups, the San, Khoi and Hadza. The Khoi are probably a creation of intermarriage between SW Bantus and San. The Hadza are an ancient group in Kenya and Ethiopia. The San form a separate race with the Somalis.
The Sandawe are another Khoisan group that was also divergent, but not enough to form a separate group, on the table here (p. 176), but was split off due to its divergence on the tree here (p. 169) .
The Sara are a a very divergent Nilotic group from Chad, who form a race with Biaka Pygmies from Central African Republic. All of the African splits are from here (p. 169).
The Funji, a Nilo-Saharan group, was both split off due to their diversity (p. 169). The Bedik, a small group of 5,000 in Senegal, are also divergent. Though they are not divergent enough to be a race on the distance chart, they are on the PC and tree charts. The Funji, or Gule, live in Sudan on the Blue Nile near the Ethiopian border (p. 170). The Bedik are a small group in Senegal.
Three groups in Senegal, the Peul, Serer (650,000) and Wolof (2 million), were split off into a separate group although they they do not have enough distance in the distance chart to warrant that, similar to the Southern Chinese, Thai and Khmer. However, like these three groups, the Senegalese groups are quite different on the PC Chart and on the tree chart, so they were split off (p. 181-182).
The Peul (700,000) speak Fulani (Peul is just French for Fulani), but are settled African farmers, unlike the more pastoralist Caucasian – Berber group that roams across the Sahel.
Figure 1 appears to divide humanity into four racial squares – Northeast Asian, Southeast Asian, Caucasian and African. Although the difference between SE and NE Asians is deeper than that between Asians and Caucasians, it is clear that this is all one race – the Mongoloids. Inside of that group, all of the Chinese are related.
The homeland of the proto-Asians dates back over 60,000 years and is in northern Vietnam and southern China. We know this because the Vietnamese have the greatest genetic diversity in all of Asia. The split between the NE Asians and the SE Asians is at least 53,000 years deep. There is a Hmong-specific line alone that may date as far back as 26,000 years.
The traditional tripartite system favored today by racial minimalists – Caucasian, Mongoloid and Negroid – is appealing, but I could not reproduce it. As there is as much difference between Asians and Caucasians as between SE Asians and NE Asians, why should I create a Mongoloid Race?
Instead, I split it into nine separate major races. This enabled me to account for the fact that while Australoids are Asians (genetic analysis of various Australoids has proven this), they are definitely an extremely divergent group.
This analysis also recognizes the deep diversity of Australoids – the Aborigines are more distant to Africans than any other race (once again despite physical appearance), due to genetic drift in Australia for millenia.
At first I put Papuans into an Australoid Race with Aborigines, but later I split them off. The distance between Aborigines and Papuans is as great as between Caucasians and Asians, so why lump the two Oceanians together? At the same time, we should recognize that there is a Mongoloid super-group that does encompass Aborigines, Papuans and both NE and SE Asians.
Figure 1 puts Aborigines barely into the NE Asian square, Papuans on the line between SE and NE Asians and Melanesians further down in the SE Asian square. Figure 4 shows that Aborigines they are mostly closely related first to Mongolians and Siberians and next to Japanese and Koreans. This is due to the Ainu substructure in these groups.
I also reluctantly split off the Kalash into a separate major race, inside of Caucasians, based on a stunning paper that differentiated the Kalash among groups such as Africans, East Asians, Oceanians, etc.
Based on Cavalli-Sforza’s six-race theory above in part, I split off Amerindians into a separate race inside of Asians. I also split off Pacific Islanders into a group called Oceanians, but contra Cavalli-Sforza, I did not include Papuans with the rest of the Pacific Islanders.
My Pacific Islander group includes Melanesians, Micronesians and Polynesians. Note that one group of Indonesians is included in each of the Melanesian and Micronesian subgroups. Therefore, there is no Indonesian race per se, as Indonesians encompass a variety of groups, although most can be put into a few SE Asian minor races.
That is based on genes. If you go by anthropometrics, you can get a group called Australoids that includes Negritos, Melanesians, the Ainu, Papuans, Aborigines, the Senoi, Tamils and Fuegian Amerindians.
The Andaman Islands Negritos are also profoundly different from other groups, and are said to have the “purest” genetic profile of any group, once again due to genetic drift and lack of outside inputs. Papuans, Melanesians and Negritos are also extremely distant from Africans, once again despite physical appearances.
The Khoisan (San and Bushmen) in Africa are the oldest race on Earth based on genetic signatures dating back 53,000 years, and this is what the original humans who came out of Africa 70,000 years ago may have looked like.
The various Negrito groups, the Aborigines and possibly the Papuans are also very ancient.
Mongoloids as we now know them are only 9,000 years old – previous groups in Asia looked more like Australoids – of which the Ainu and Gilyak are the last remaining descendants.
Australoid types and their ancestors are the original peoples of India , Burma, Thailand, Vietnam, Cambodia, Philippines, Indonesia, and possibly even New Guinea and Australia. For instance, the Semang go back an incredible 50,000 years in Malaysia.
The Bantu (or the Africans that we are familiar with) may go back much further – it has been up to 40,000 years since they split off from the Pygmies. There is a suggestion that they were distinguishable from Khoisan (Bushmen) even 100,000 years ago (p. 160). The ancestors of all Africans seem to have come from West Africa at least 35,000 years ago (p. 160).
Amerindians at the tip of South America are very different in head shape than the rest of the Amerindians – looking more like Australoids – and their genetics is also profoundly different.
The proto-Caucasian homeland may have been in the Caucasus about 45,000 years ago. Another theory says it was in Central Asia.
The most ancient Europeans are the Saami and an ancient, isolated group of Sardinians. Among Caucasians, the Berber and South Indian Races appear to be very ancient, and both are extremely divergent within the Caucasian group. They may be surviving remnants of the most ancient Caucasians.
The South Indians are actually midway between Caucasians and Asians genetically and are only lumped with Caucasians because this is who they most resemble.
Europeans proper only go back 10,000 years or so, but the Saami (best seen as proto-Europeans) seem to go further back than that.
South Indians have been evolving in considerable isolation for about 15-20,000 years in the subcontinent. Prior to that, they appear to have come from the Middle East. The Berbers of today appear to be continuous with Berbers of up to 50,000 years ago, making them the most ancient Caucasian race of all.
The rest of the groupings mostly follow from Figure 1. More tables like Table 4 in Capelli would be very helpful in order to tease out more minor races.
A single asterisk indicates considerable genetic difference from related groups, two asterisks indicates a highly divergent group, and three asterisks is a profoundly divergent group. Major races are in red.
Some groups are not represented. I was not able to classify many groups with Negrito or Veddoid affiliations, such as the Tamils of South Asia and the Montagnards of Vietnam.
Mien and Qiang are Northern Chinese tribes, but the Mien have moved to the South lately. I could not find any good genetic data on the Qiang. The Nu were arbitrarily included in the Tibetan Race because they came from Tibet, but I don’t have good genetic data to prove that this is really a single unit. The chart here does not clarify things much.
The Bhutanese, though most closely related to Tibetans, were given their own race based on data showing that they are nevertheless considerably distant from Tibetans.
The Barya are a mixed-race group in Western Eritrea.
The Gilyak or Nivkhi are an ancient tribe living on the border between Korea, Russia and Japan that has ties to the Ainu. Ryukyuan is another name for Okinawan. They were given a separate race based on studies showing them intermediate between the Ainu and modern Japanese.
The Va (or Wa) are an ethnic group in Yunnan and Burma that seems to be distinct from the Northern, Southern and Tibetan Chinese groups. The Va seem to be about equally related to the Northern and Southern Chinese, indicating some sort of a dual origin. The Jingpo, or Karen, another Yunnan group that also occurs in Burma, were included with them based on this paper. The Lawa of Thailand were added to this group based on Figure 5. Interestingly, the languages of the Lawa and Va are also closely related.
A Southern Japanese Race was split off from the Japanese, Ryukuyans and Ainu. This group is made up of Kyushu Island, the southernmost island, and the Kinki region of Honshu, near the city of Kyoto. The Japanese in this area are highly divergent (p. 232).
The European-Iranian Race includes almost all Europeans except the Saami, Basques and Sardinians. The Saami and the Sardinians are very distant and the Basques much less so from the rest of the Europeans.
Although Cavalli-Sforza classes the Basques, Yugoslavs and Greeks as genetic outliers, there was not enough distance between the Yugoslavs and Greeks and other Europeans to split them into a separate group on the basis of genetic distance. Furthermore, the Greeks are clearly in the European group in Fig. 1 – they are quite close to English and Danes in the PC analysis.
However, I did split the Basques off based on their lying outside the European-Iranian cluster on the PC chart in Fig. 1. Most groups that were distinguished as independent units outside of clusters on Fig. 1 were given separate races.
The Greeks are interesting in that, while they are obviously a part of the Europeans on all charts, they are also the only Europeans that are are also close enough to most Middle Easterners to be included in their group. So the Greeks are a link between the European and Middle Eastern groupings inside the Caucasian Race.
The Iranian branch includes Jordanians, Iraqis, Assyrians, Druse, Lebanese, Kurds, Georgians, Caspians, Turks, Jews, and related groups in the area. It was difficult to decide whether to put the Turks in the Iranian subgroup or in the Central Asian subgroup, as they are close to both.
It was also very difficult to decide whether to put the people of the Caucasus, the Kurds, Turks, Caspians and Jews in the Iranian group or the Central Asian group as they cluster with both. I decided on sheer geographic grounds to put them in the Iranian group. The Russian Saami are closer to the Tungus and were included in that group.
Although some Arabs, West Asians and all South Indians were split off, this was somewhat arbitrary. Although they form separate groups on the Fig. 1, the Arabs are closely enough related to various Europeans, including Greeks, to be included with Europeans (Fig. 4). However, the Arabs were not as close as the Iranians.
Likewise, South Indians are close to Iranians, who are in turn close to Greeks and Italians – note that Iranians are also somewhat close to Danes and English (Fig. 4). As the Greeks link Europeans genetically with Middle Easterners, the Iranians link Europeans genetically with India. Arabs and South Indians were only split off due to the distance observable in Fig. 1.
West Asians were also split off due to their divergence. Based on this chart, they seem to be a compact grouping. This group includes the Pashtuns, Brahuis, Balochis, Makranis and Sindhis.
Further research shows that the Tajiks and Hunza, who at first appear to group with the West Asian group above, actually compose two groups divergent enough to be split into 2 different races. The first group is made of the Hunza of the Karokorams, the Bartangi of the Pamir Range and the Roma or Gypsies of Europe. So the Gypsies have a Himalayan origin.
The second group is made up of Tajiks, the Shugnan of the Pamirs, Bukhara Arabs and three groups in India – the Kallar of Kerala, the Sourashtran of Tamil Nadu and Yadhava of various parts of the region.
The Kalash, a strange, ancient, tiny tribe with Caucasian roots in northwest Pakistan in Chitral Province, are so diverse that they could very well form their own major grouping entirely, on a par with Africans, Europeans – Middle Easterners – West and South Asians, Oceanians, East Asians and Amerindians.
Since making a macro race out of a tiny ethnic group in Pakistan is absurd, I decided to throw them as a major race subsumed under Caucasians, albeit on the grounds that they are an extremely divergent race. They were classed with Caucasians because there is a general consensus that this is what they are (last two links are racist).
Due to their divergence, Kuwaitis and Arabians – consisting of Saudis, Yemenis and Bedouins – were split off into separate groups.
The are numerous groups that are more or less recent combinations of various groups and do not yet deserve their own racial category.
Hispanics are in general a mixture between Caucasians (typically Iberians) and Amerindians. They have been evolving for a short time and have not had time to differentiate into anything suggesting a race yet (despite nonsense from La Raza demagogues).
There are other Hispanics who are heavily mixed with Blacks, Caucasians and Amerindians. This is especially seen in South America in Brazil, Venezuela, and Colombia, and even in Central America and Mexico.
There are large Black-White mixed populations in the West Indies. In Singapore and Hawaii, there are rapidly mixing populations that defy categorization.
This paper is basically just a shot in the dark and is more properly termed a pilot or exploratory study. I welcome evidence-based inputs from any knowledgeable persons who wish to add to this preliminary grouping of the human races, major and minor. All suggestions coming from nationalists of various types, ethnic or otherwise, typically lacking evidence, will probably be rejected outright.
There are 4 macro races of man, 11 major races of man and 115 minor human races of man.
* = significant genetic distance from most other groups
** = major genetic distance from most other groups
*** = extreme genetic distance from most other groups
Asian Macro Race
Northeast Asian Major Race*
Japanese-Korean Race (Japanese – Korean)
Southern Japanese Race (Honshu Kinki – Kyushu)
Ryukyuan Race (Okinawans)
Ainu Race*** (Ainu)
Gilyak Race** (Gilyak)
Northern Chinese Race (Northern Han – Qiang – Manchu – Hui – Yunnan Han)
Oroqen Race (Oroqen)
Sherpa-Yakut Race (Sherpa – Yakut)
Nepalese Race (Nepali – Newari)
Mongolian Race (Mongolian – Inner Mongolian – Buryat – Kazakh)
Northern Turkic Race*** (Dolgan – Altai – Shor – Tofalar – Uighur – Chelkan – Soyot – Kumandin Teleut – Hazara)
Central Asian Race (Kirghiz – Karalkalpak – Uzbek – Turkmen)
Tuva Race (Tuva)
Tungus Race (Even – Evenki – Russian Saami)
Siberian Race
Beringian Race** (Chukchi – Aleut – Siberian Eskimo)
Paleosiberian Race (Koryak – Itelmen)
Reindeer Chukchi Race (Reindeer Chukchi)
General Tibetan Race (Tibetan – Lisu – Nu – Tujia – Akha – Burmese – Yizu)
Mizo Race (Mizo)
Bhutanese Race (Bhutanese Buddhist)
Siberian Uralic Race (Nentsy – Samoyed – Ket – Mansi – Khanty)
Nganasan Race (Nganasan)
Uralic Race (Komi – Mari)
North American Eskimo Race (Inuit)
Amerindian Major Race*
Northern Na-Dene Race
Northwestern American Amerindian Race
Northern Amerind Race
Central Amerind Race
Southern Amerind Race
Ge Amerindian Race (Ge Language Group)
Tucanoan Amerindian Race (Tucanoan Language Group)
Nootka Amerindian Race (Nuuchahnulth – Makah)
Fuegian Amerindian Race (Ona – Yaghan – Kaweskar – Aonikenk – Alacaluf)
Southeast Asian Major Race*
Southern Chinese Race (Dong – Henan Han – Yi – She – Punu – Naxi)
Hmong-Mien Race (Chinese Hmong – Thai Hmong – Mien)
Li-Khmer Race (Li – Khmer)
Southeast China Race (Hakka – Min Nan – Singapore Chinese – Thai Chinese – Cantonese Han)
South China Sea Race (Tagalog – Ilocano – Visayan – Ami Taiwanese Aborigine – Guangdong Han)
Manobo Race (Manobo)
Philippines Negrito Race (Aeta – Agta – Palau Micronesian)
Mangyan-Ati Race (Iraya – Ati)
Mamanwa Philippines Negrito Race (Mamanwa)
Tai Race (Thai – Tai Lue – Tai Kern – Tai Yong – Tai Yuan – Lao – Lahu – Aini – Shan – Dai – Muong – Buyei)
Vietnamese Race (Vietnamese – Deang – Jinuo – Blang)
Mlabri Race** (Mlabri)
Htin Race (Htin)
Kachin Race (Kachin – Karen – Va – Nung – Lu – Lawa)
General Taiwanese Aborigine Race (Ayatal – Bunun – Yami)
Island SE Asian Race (Paiwan Taiwanese Aborigine – Sea Dayak – Sumatran – Balinese)
Bidayuh Race** (Jagoi)
Indonesian Race (Sulawesi – Borneo – Lesser Sunda – Sarawak – Javanese)
Mentawi Race (Mentawi)
Toraja Race (Toraja)
Lesser Sunda Race (Kambera – Lembata – Lamaholot – Manggarai)
Malay Race (Malaysia Malay – Singapore Malay)
Proto-Malay Race** (Temuan)
Austroasiatic Race (Mon – Zhuang – She – Ho – Lyngngam)
Nongtrai Race (Nongtrai)
Santhal-Naga Race (Santhal – Naga – Munda – Kurmi – Sudra)
Meghalaya Race (War Jantia – Bhoi – Maram – War Khasi – Kynriam – Nishi – Pnar – Bai)
Senoi Race (Senoi)
Shompen Race (Shompen)
Garo Race (Garo)
NE Indian Indo-European Race (Mahishya – Bagdi – Gaud – Tanti – Lodha)
Indian Tibeto-Burman Race (Apatani – Nishi – Adi – Tripuri – Jamatia – Mog – Chakma)
Semang Malay Negrito Race*** (Semang – Jehai – Kensui)
Oceanian Major Race*
Micronesian Race (Yap – Kanaka – Toba Batak Indonesian – Kora Batak Indonesian)
Polynesian Race* (Tonga – Western Samoa – French Polynesia – Cook Islands)
Melanesian Race (Fiji – Vanuatu – New Ireland – Papuan Melanesian – Nasioi – Alor Indonesian)
Australoid Macro Race
Australian Major Race***
General Australian Aborigine Major Race***
Queensland Aborigine Race***
Western Territory Pama-Nguyan Aborigine Race***
Papuan Major Race***
General Papuan Race***
Motu Papuan Race***
Sepik-Ramu Papuan Race***
Greater Andaman Islands Major Race***
Greater Andaman Islands Negrito Race***
Onge Andaman Islands Major Race***
Onge Andaman Islands Negrito Race***
Caucasian Macro Race
General Caucasian Major Race***
European-Iranian Race (Most European – Caucasus – Armenian – Jewish – Turk – Kurd – Iranian – Jordanian – Iraqi – Assyrian – Druze – Lebanese – Georgian – Caspian – Palestinian)
Basque Race (Basque)
Norwegian-Swedish Saami Race*** (Norwegian Saami – Swedish Saami)
Finnish Saami Race** (Finnish Saami)
Sardinian Race** (Sardinian)
Kuwaiti Race* (Kuwaiti)
Arabian Race (Saudi – Yemeni – Bedouin)*
West Asian Race (Pashtun – Brahui – Balochi – Makrani – Sindhi )
Tajik Race (Tajik – Bukhara Arab – Shugnan – Kallar – Sourashtran – Yadhava)
West Himalayan Race (Hunza – Bartangi – Roma)
Berber Race*** (Berber)
Egyptian Race (Egyptian)
North African Race (Moroccan – Libyan – Tunisian – Canarian)
Algerian Race (Algerian)
North Indian Race** (Punjabi – Central Indic – Punjabi Brahmin – Rajput – Vania Soni – Mumbai Brahmin – Jat – Kerala Brahmin – Koli)
Himalayan Race*** (Gurkha – Tharu – Ladakhi)
Karnet-Uttar Pradesh Brahmin Race*** (Karnet – Uttar Pradesh Brahmin)
South Indian Race** (Munda – Bhil – Maratha – Rajbanshi – Oraon – Parji – Kolami Naiki – Chenchu Reddi – Konda – Kolya – West Bengal Brahmin – Parsi – Gond)
Kerala Kadar Race*** (Kerala Kadar)
South Dravidian Race*** (Sinhalese – Lambada – Irula – Izhava – Kurumba – Nayar – Toda – Kota – Malayaraya – Tamil)
Kalash Major Race***
Kalash Race*** (Kalash)
African Macro Race
African Major Race***
Tigrean Race*** (Tigrean)
Amharic Race*** (Amharic)
Sudanese-Barya Race*** (Sudanese – Barya)
General Nilotic Race (Shilluk – Masai – Nuer – Dinka – Luo – Turkana – Karanojo – Mabaan)
Funji Nilotic Race (Funji)
Tuareg-Beja Cushitic Race*** (Tuareg – Beja)
Nubian Race*** (Nubian)
Wolof-Peul-Serer Race (Wolof – Peul – Serer)
General Bantu Race (Most Bantus)
Bedik Bantu Race (Bedik)
West African Race (Most West Africans)
Mbuti Pygmy Race
Sara Nilotic-Biaka Pygmy Race (Sara – Biaka)
San Khoisan-Somali Race*** (San – Somali)
Khoi Khoisan Race*** (Nama – !Ora)
Hadza Khoisan Race*** (Hadza)
Sandawe Khoisan Race (Sandawe)
References
-
Capelli C., Wilson J. F., Richards M., Stumpf M. P. H., Gratrix F., Oppenheimer S., Underhill P., Pascali V. L., Ko T. M., and Goldstein D. B. 2001. A Predominantly Indigenous Paternal Heritage for the Austronesian-Speaking Peoples of Insular Southeast Asia and Oceania. American Journal of Human Genetics 68:432-443.
Cavalli-Sforza L. L., Menozzi P,. Piazza A.. 1994. The History and Geography of Human Genes. Princeton, NJ: Princeton University Press.
Chu J. Y., Huang W., Kuang S. Q., Wang J. M., Xu J. J., Chu Z. T., Yang Z. Q., Lin K. Q., Li P., Wu M., Geng Z. C., Tan C. C., Du R. F., and Jin L.. 1998. Genetic Relationship of Populations in China. Proceedings of the National Academy of the Sciences of the United States of America (PNAS). 95:11763-11768.
Harihara S., Saitou N., Hirai M., Gojobori T., Park K. S., Misawa S., Ellepola S. B., Ishida T. and Omoto K. 1988. Mitochondrial DNA Polymorphism Among Five Asian Populations. Am. J. Hum. Genet. 43:134-143
Jablonski, N. and Chaplin, G. 2000. The Evolution of Human Skin Coloration. Journal of Human Evolution. Available on this blog here.
Lĭ H., Pan S., Donnelly M., Tran D., Qin Z., Zhang Y., Cheng X., Yin R., Lin W. and Hoang V. 2006. Dermatoglyph Groups Kinh Vietnamese to Mon-Khmer. International Journal Of Anthropology 21:3-4, pages 295-306.
Lin M, Chu CC, Chang SL, Lee HL, Loo JH, Akaza T, Juji T, Ohashi J, Tokunaga K. March 2001. The Origin of Minnan & Hakka, the So-called “Taiwanese”, Inferred by HLA Study. Tissue Antigens:57(3):192-9.
Omoto, K. (1984). The Negritos: Genetic Origins and Microevolution. Acta Anthropogenetics 8(1-2):137-47.
Omoto K., Ueda S., Goriki K., Takahashi N., Misawa S., and Pagaran I. G. (1981). Population Genetic Studies of the Philippine Negritos. III. Identification of the Carbonic Anhydrase-1 Variant With CA1 Guam. Am J Hum Genet. 33(1): 105-111.
Reddy BM, Langstieh BT, Kumar V, Nagaraja T, Reddy ANS, et al. 2007. Austro-Asiatic Tribes of Northeast India Provide Hitherto Missing Genetic Link Between South and Southeast Asia. PLoS ONE 2(11): e1141.
Useem, John. 1948. Human Resources of Micronesia. Far Eastern Survey, Vol. 17, No. 1. pp. 1-4.
